# Principal Engineer, AI Security at Lila Sciences

- Company: Lila Sciences
- What the company does: LILA has created the world's first Operating System for Science powered by Scientific Superintelligence™. Backed by General Catalyst.
- Company website: https://www.lila.ai/
- Type: Startups (AI role)
- Level: Principal and up
- Location: Cambridge, MA USA
- Work setup: On-site
- Posted: 2026-10-01
- Apply by: 2026-11-23
- Apply: https://job-boards.greenhouse.io/lilasciences/jobs/4403714009?utm_source=1752vc&utm_medium=careers
- Page: https://www.1752.vc/careers/jobs/lila-sciences-principal-engineer-ai-security/

## About the role

Lila Sciences is building autonomous systems for scientific discovery, and the AI infrastructure behind that work needs security built in from the start. This role sits on the IT & Security team and owns security engineering for Lila's AI and ML systems, from training and inference pipelines to model registries, MCP servers, and agent runtimes. Found on 1752vc Careers, the job board for startup and VC roles.

## What they're looking for

- Strong background in security engineering, cloud security, or DevSecOps, with hands-on experience securing production infrastructure
- Working knowledge of AI/ML systems and the ML lifecycle, plus AI-specific risks such as prompt injection, model and data poisoning, insecure agent tooling, and inference-time threats
- Experience with software supply chain security and dependency and vulnerability management
- Experience implementing data security controls (classification, data flow mapping, access control, encryption) in cloud environments
- Proven ability to partner across engineering teams and drive cross-functional remediation to closure
- Comfort operating with ambiguity in a fast-moving research environment

Tags: Business Operations

Source: 1752vc Careers, https://www.1752.vc/careers/jobs/lila-sciences-principal-engineer-ai-security/